Greater Yellowstone's grizzly population is the best known in the world and one of the symbols of its beloved status is 24-year-old Jackson Hole grizzly mother 399 who this spring emerged from her den with four cubs This image, titled "Among the Sage—399 and Quadruplets" is a new print release by Thomas Mangelsen and we are grateful for his permission to use it. You can view more of Mangelsen's work at Mangelsen.com